In addition, the assumption that a ram by a 30-60 ton whale would be just a sort of minor nuisance that might temporarily deter the shark is frankly wrong.
Even orcas (and reportedly even bottlenose dolphins) use rams to lethal effect, even against larger whales (and sharks), and despite being themselves an order of magnitude smaller than
Livyatan. In these taxa, damage seems to be primarily localized, which explains why they don’t require a large dampening spermaceti organ, but a pointed, stiff rostrum. Physeteroids however are a different story.
The ramming adaptations and capabilities of extant
Physeter have been studied. At the scales we are talking about, a single ram at reasonably high speed may well end the fight (and not just temporarily) or at least cause major injury, simply by inducing a fatal level of accelleration on even a large-bodied opponent.
Carrier et al. 2002 simulated that under all their assumptions for dampening constants, a
Physeter hit by another
Physeter at 3m/s (about 11 km/h) would experience g-forces at which fatal injuries would occur, despite only mild accellerations to the attacking whale. Now as I am sure someone will bring up,
Livyatan does have a smaller spermaceti organ than
Physeter, but it is still seriously sized, I remember the head alone coming out at over 9 t in my model, and that was under very conservative assumptions regarding the size of the cranial soft tissues. The giant size of the spermaceti organ (and junk) in sperm whales is also subject to other pressures than just the one towards ramming, and hence might be overbuilt for this purpose alone.
And we are not requiring the
Livyatan to necessarily kill the shark in one single ram or be able to sink a whaling ship of over 200 tons, it is enough to cause serious injuries (not something the shark would just take as a cue to retreat and come back later, something that would be life threatening, at the very least while faced with a large, formidable opponent). Which we can even expect to happen more easily to a shark than another whale, as we also need to consider the local effects the concentration of such forces on a relatively small area (the frontal area of the whale‘s junk) could have against some of the not so well protected parts of the shark (chondocranium, gills, internal organs…).
This is not a mildly unpleasant push that might cause the shark to back off and come back later. We are talking about a kinetic energy at least equivalent to being hit by a bus. Essentially, a battering ram. The physeteroid head has adaptations specifically to withstand and dissipate this energy (Panagiotopoulou et al. 2016). The shark simply does not. A head on collision with a 30-60 ton whale? From what we know, potentially fatal.
It seems periodic reminders of this are needed, as no matter how often the evidence is posted, after a while people revert back to thinking of ramming as some sort of mild annoyance.
This seems to be a one-sided perspective.
Majority of odontocetes including delphinids exhibit ramming behavior for different ends including as a defensive measure.
Even some of the mysticetes are known to exhibit ramming behavior at times; Gray whales and Humpback whales are usual suspects in modern times.
Cetaceans at large, are also known to employ tail slapping technique to defend themselves at times.
Therefore, sharks which are attempting to hunt cetaceans, are likely to experience ramming as well as tail slapping at some point. How many sharks have succumbed to these blows by the way? Any statistical data?
POINT OF CONTENTION A macropredator which clearly evolved for
big-game hunting, must have necessary biological adaptations to cope with affiliated risks and/or pressures, or it will not be up to the task.
Megalodon did not had a purposeless existence; it had a profound impact on ecosystems of its time. Megalodon was the greatest natural agent to ensure checks and balances on the populations of cetaceans across the board including in the case of
Livyatan.
So what kind of biological adaptations would be necessary to enable
big-game hunting for sharks in marine environments? Some of these are listed below:
- Gigantism (Wroe et al., 2008);
- Excellent killing apparatus (Wroe et al., 2008);
- Regional endothermy (Ferrón, 2017);
- Thickness of cartilage*
MERITS OF CARTILAGEFor starters, cartilage have superior shock-absorbing properties as well as elasticity than solid bones.
Why do you think human knees contain cartilage? Knees are subject to much higher amount of stresses than other parts of the body during physical activities and solid bones are not up to the task, so cartilage enable humans to perform functions and cope with affiliated pressures which they cannot otherwise.
Enlightening study for reference:-
Malekipour, F., & Lee, P. V. (2019). Shock absorbing ability in healthy and damaged cartilage-bone under high-rate compression. Journal of the mechanical behavior of biomedical materials, 90, 388-394.
Key observation is that
thickness of cartilage will make substantial difference in terms of coping with extreme forces (Malekipour & Lee, 2019), and therefore an important consideration.
*
NOT ALL SHARKS ARE CREATED EQUALWhen WE take a look at fossilized remains of Megalodon, we notice cartilaginous structures that are extremely ROBUST/STRONG on average, or used to be in real life.
Closer look
Megalodon's cartilaginous structures put that of others to shame in all aspects. WE not talking about just a shark here, WE are talking about
the shark here.
Megalodon's cartilaginous structures are/were clearly built to cope with extreme levels of stresses as well as be excellent in 'shock-absorbing' capacity. This biological characteristic is in line with adaptations to cope with the stresses of
big-game hunting in marine environments.
Megalodon was likely to experience ramming as well as tail slapping from cetaceans on a frequent basis, and it had to be strong enough to endure these blows when they came. This shark should be able to take a lot of punishment in its trophic interactions because it doesn't have the luxury to slip away from a situation on a moment's notice; this shark is/was a multitonned macropredator (in the higher end of the spectrum) after all.
Unfortunately, little attention is paid to biomechanical considerations for Megalodon, how it was coping with competitive pressures in various environments, and what kind of competitive pressures it was subjecting other lifeforms to in turn. WE need studies like Wroe et al (2008), Ferrón (2017) and Gilbert et al (2018) to understand the bigger picture here. Megalodon's biological capabilities and resultant 'competitive game' can be far better understood from biomechanical considerations and viewpoints.
Much of the focus is directed towards pointless discussions about when Megalodon became extinct and which animals were 'supposedly' responsible. Talk about misplaced priorities and being excessively dumb. And I am rather worried that people are being awarded PhDs for this kind of stupidity.
FORCES IN QUESTIONGiven Megalodon's physiology, it would require something substantially impactful to drive it away. However, few would be up to the task, to produce forces of such magnitude.
Livyatan melvillei is a notable candidate of-course.
Life 1) The site that sold this tooth claims it is
Physeter sp.
paleodirect.com/wh023-largest-complete-fossil-sperm-whale-tooth-with-megalodon-shark-bites/It’s not that I particulatly trust them (they also have a rather colorful interpretation on how this happened, "fight with the shark and whale locking jaws"), but the tooth does look similar to
Physeter teeth, they provide a figure plate of a
Physeter skull and teeth showing this:
paleodirect.com/product_images/uploaded_images/odont1.jpgIt could also be a
Livyatan tooth, anatomically that would be plausible as well. In that case, the first from the northern hemisphere.
Anyway, what they interpret quite confidently as a megalodon bite mark also seems a bit overinterpreted. Even is it is though, how is that evidence megalodon preyed on this whale? A bite to the jaw would appear a very strange predation technique indeed.
2) Of course. I would presume either one would want to end the fight quickly, but of course the question is if they could, i.e. if their opponent would let them do this. For that reason, of course it would seem exceedingly unlikely they would really fight on equal ground in a natural setting, even one preying on the other with the benefit of ambush would seem rather unlikely due to the risk involved.
4) I am not doubting this. Surely, megalodon was one of the greatest macropredators to have ever lived. As was
Livyatan.
5) thanks. Yes, I know the article. As I recall, that is a piece of popular science authored by a journalist, not by Bretton Kent.
1. Physeter sp. = unknown species / undetermined species
"This is a MASSIVE and impressive specimen - the BEST we have ever handled or offered. It is a LARGE, complete tooth from a prehistoric Sperm Whale of the family Physeteridae dating back to the Miocene/ Pliocene Period. Like T. rex, the Sperm Whale ruled the ancient seas and shared the title of "king" with the Megalodon Shark during its day."Now, take another look:-
That jaw structure belong to a modern-era
Physeter macrocephalus, and its teeth are similar to
Type D in the photo below.
Relevant details in this link:
theworldofanimals.proboards.com/post/47686The tooth being held in the hands with the intent to compare with others, is similar to
Type C in the
Livyatan holotype.
The tooth containing cuts from Megalodon (WH023), is also similar to
Type C in the
Livyatan holotype.
This tooth seems to indicate presence of
Livyatan types as far as in the lower North American environments. Although these fossils are rare which is strange.
WH023 = Savannah Basin (Georgia)
Do you find bite marks on this specimen unsettling?
Intent behind attacking the skull of an animal would be to kill it swiftly. Megalodon was well-equipped to cut through the bones (mechanical advantage of having a robust dentition). Now combine this quality with gigantism, and Megalodon could attack a large cetacean in numerous ways - a luxury that other sharks do not have. However, killing a large cetacean swiftly is a sound predation strategy, and also a meaningful indicator of Megalodon's intelligence and/or learning capacity.
Fossilized remains of an unknown baleen whale, excavated from Sharktooth Hill, also suggest an attack on the skull.
"A fossilized whale was also found at the quarry — missing its head. The owners of Shark Tooth Hill, Sean and Lisa Tohill, have long wondered if the whale was killed by a megalodon."Link:
www.businessinsider.com/scary-truth-about-megaladon-eating-whales-2013-8This excavation motivated development of a special episode for Shark Week TV series (Sharkzilla) in which dry-skull method was employed to understand what a Megalodon could do to other objects.
Nevertheless, WH023 suggest that large macropredatory sperm whales, or raptorial odontocetes in general, were not immune to predation from Megalodon. This is a significant discovery from which numerous inferences can be drawn which in turn shall advance our understanding of the unforgiving Miocene marine environments.
2. That the Megalodon tore through the skull a large sperm whale (WH023) in a trophic interaction, is a significant pointer towards what could happen to
Livyatan in this fight.
There is much about these two animals which WE do not understand at present, there are aspects of animals which do not fossilize.
It is difficult to determine whether Megalodon were attacking raptorial odontocetes on a regular basis (or not). Although Megalodon's demise seem to coincide with a major drop in the diversity of the Order
Physeteriodae by the end of Miocene.
Abstract of an interesting study:-
"Physeteroidea (sperm whales) attained great diversity during the Miocene and early Pliocene, and the phosphatic sands of the U.S. Atlantic Coastal Plain have produced thousands of specimens. Although postcranial and cranial materials are rare, teeth are remarkably common and have the potential to provide valuable insight into the lives of these animals. We examine a suite of Physeteroidea indet. teeth from the Lee Creek Mine to better constrain the life history and ecology of this extinct group. Wear facets indicate that these animals, unlike modern sperm whales, had both maxillary and mandibular teeth, suggesting a raptorial feeding ecology more akin to killer whales. A relationship between tooth diameter and body size established for modern odontocetes suggests that these animals as adults were also about the size of modern killer whales. Because physeteroid teeth grow continuously over ontogeny and are not replaced, counts of accretionary growth layer groups can be used to ascertain the age of an animal at death. Tallies of growth increments from 10 teeth, including some of the largest available, reveal that life spans only rarely exceeded 20 years, significantly shorter than the 65+ years typical of modern sperm whales or orcas. Despite their large size, these odontocetes experienced a ‘fast’ life history, more like beluga whales today. We suggest that the rapid growth and short life span exhibited by the Lee Creek physeteroids are, like the modern beluga, evolutionary responses to high predation pressure imposed by large co-occurring predatory taxa, particularly including Carcharocles megalodon or other large macroraptorial physeteorids.
"
From Gilbert et al (2018); citation below in the REFERENCES section.
Relatively longer lasting members of the Order
Physeteriodae were probably already specialized predators of squids and small fish, or slowly changed form (evolved) to hunt squids and small fish in order to better cope with the pressures of Pliocene (cooling trends; geographical shifts; mass extinction events).
3. Nothing in your response so addendum.
4. WE need more insight about
Livyatan's dietary preferences and not jump to conclusions on the basis of superficial observations. I notice this
bias even in professional circles although many underestimate Megalodon in their estimations which is plainly stupid.
5. Not an issue because
Hell's teeth contain valuable insight from professionals, just not articulated like a formal scientific publication (unfortunately), but good enough to refer to when necessary.
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REFERENCESFerrón, H. G. (2017). Regional endothermy as a trigger for gigantism in some extinct macropredatory sharks. PloS one, 12(9), e0185185.
Gilbert, K. N., Ivany, L. C., & Uhen, M. D. (2018). Living fast and dying young: life history and ecology of a Neogene sperm whale. Journal of Vertebrate Paleontology, 38(2), e1439038.
Wroe, S., Huber, D. R., Lowry, M., McHenry, C., Moreno, K., Clausen, P., ... & Summers, A. P. (2008). Three‐dimensional computer analysis of white shark jaw mechanics: how hard can a great white bite?. Journal of Zoology, 276(4), 336-342.
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